1. Faculty of Life and Environmental Sciences, University of Tsukuba , Tsukuba , Japan
2. Department of Biology, School of Arts and Sciences, National University of Mongolia , Ulaanbaatar , Mongolia
3. Botanic Garden and Research Institute, Mongolian Academy of Science , Ulaanbaatar , Mongolia
4. Graduate School of Environmental Studies, Nagoya University , Nagoya , Japan
5. Arid Land Research Center, Tottori University , Tottori , Japan