1. Aging Institute of University of Pittsburgh and University of Pittsburgh Medical Center , Pittsburgh, PA 15219, USA
2. Division of Cardiology, Department of Medicine, University of Pittsburgh School of Medicine , Pittsburgh, PA 15219, USA
3. Proteases and Tissue Remodeling Section, National Institute of Dental and Craniofacial Research, National Institutes of Health , Bethesda, MD 20892, USA
4. Microbial Pathogenesis Section, Laboratory of Parasitic Diseases, National Institute of Allergy and Infectious Diseases, National Institutes of Health , Bethesda, MD 20892, USA
5. Division of Infectious Diseases, Department of Medicine, University of Pittsburgh School of Medicine , Pittsburgh, PA 15219, USA