1. Division of Immunology and Genome Biology, Medical Institute of Bioregulation, Kyushu University , 3-1-1 Maidashi, Higashi-ku, Fukuoka 812-8582 , Japan
2. Department of Otorhinolaryngology, Graduate School of Medical Sciences, Kyushu University , 3-1-1 Maidashi, Higashi-ku, Fukuoka 812-8582 , Japan
3. Division of Transcriptomics, Medical Institute of Bioregulation, Kyushu University , 3-1-1 Maidashi, Higashi-ku, Fukuoka 812-8582 , Japan
4. The Walter and Eliza Hall Institute of Medical Research , Parkville, VIC 3050 , Australia
5. Department of Medical Biology, The University of Melbourne , Parkville, VIC 3010 , Australia