1. Institute of Agrobiological Sciences, National Agriculture and Food Research Organization , 1-2 Owashi, Tsukuba, Ibaraki 305-0851 , Japan
2. Faculty of Science, Academic Assembly, Toyama University , 3190 Gofuku, Toyama 930-8555 , Japan
3. Graduate School of Agricultural and Life Sciences, University of Tokyo , 1-1-1 Yayoi, Bunkyo-ku, Tokyo 113-8657 , Japan
4. Faculty of Agriculture, Setsunan University , 45-1 Nagaotogecho, Hirakata, Osaka 573-0101 , Japan