1. Department of Infection Control Science, Meiji Pharmaceutical University , 2-522-1 Noshio, Kiyose, Tokyo 204-8588 , Japan
2. Laboratory of Microbiology and Infection Control, Kyoto Pharmaceutical University , 5 Misasagi-Nakauchi-cho, Yamashina-ku, Kyoto 607-8414 , Japan
3. Department of Intractable Diseases, National Center for Global Health and Medicine , 1-21-1 Toyama, Shinjyuku-ku, Tokyo 162-8655 , Japan
4. Pathogenic Microbe Laboratory, Research Institute, National Center for Global Health and Medicine , 1-21-1 Toyama, Shinjyuku-ku, Tokyo 162-8655 , Japan