1. Department of Ecology, Environment and Plant Sciences, Stockholm University, Stockholm, Sweden
2. Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark
3. Department of Botany, University of British Columbia, Vancouver, British Columbia, Canada
4. UBC Botanical Garden & Centre for Plant Research, University of British Columbia, Vancouver, British Columbia, Canada
5. Naturalis Biodiversity Centre, Leiden, The Netherlands
6. Department of Environmental Science, Aarhus University, Denmark