1. Chinese Academy of Sciences (CAS) Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization & Ecological Restoration and Biodiversity Conservation Key Laboratory of Sichuan Province, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, China
2. College of Life Sciences, Shanghai Normal University, Shanghai, China
3. Department of Biology, Terrestrial Ecology Section, University of Copenhagen, Copenhagen, Denmark
4. Center for Permafrost (CENPERM), University of Copenhagen, Copenhagen, Denmark