1. CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla, Yunnan 666303, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla, Yunnan 666303, China
4. Horticulture Department, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla, Yunnan 666303, China
5. Yuanjiang Savanna Ecosystem Research Station, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Yuanjiang, Yunnan 653300, China