1. Department of Sustainable Land Management and Soil Research Centre, School of Agricultural Policy and Development, University of Reading , Reading, Berkshire RG6 6AR , UK
2. School of Biological Sciences, The University of Western Australia , 35 Stirling Highway, Crawley, WA 6009 , Australia
3. School of Agriculture and Environment, The University of Western Australia , 35 Stirling Highway, Crawley, WA 6009 , Australia