1. Research Centre for Applied Alpine Ecology, La Trobe University, Bundoora, VIC 3086, Australia
2. Department of Ecology, Environment and Evolution, La Trobe University, Bundoora, VIC 3086, Australia
3. Centre for Integrative Ecology, Deakin University, Burwood, VIC 3125, Australia
4. Centre of Excellence for Biosecurity Risk Analysis, The University of Melbourne, Parkville, VIC 3010, Australia