1. School of Biological Sciences, University of Auckland , Auckland , New Zealand
2. Department of Zoology, University of Otago , Dunedin , New Zealand
3. Institute of Marine Science, Leigh Marine Laboratory, University of Auckland , Warkworth , New Zealand
4. Institut de Systématique, Évolution, Biodiversité (ISYEB), École Pratique des Hautes Études | PSL, MNHN, CNRS, Sorbonne Université, Université des Antilles , Paris , France