1. X-STAR, Japan Agency for Marine-Earth Science and Technology (JAMSTEC) , Yokosuka, Kanagawa , Japan
2. Univ Brest, Ifremer, CNRS, Unité Biologie des Environnements Extrêmes marins Profonds , Plouzané , France
3. Ocean and Earth Science, National Oceanography Centre Southampton, University of Southampton , Southampton , United Kingdom