1. Instituto Gulbenkian de Ciência , Oeiras , Portugal
2. Laboratory of Genetics, University of Wisconsin-Madison , Madison, WI , USA
3. cE3c - Centre for Ecology, Evolution and Environmental Changes & CHANGE - Global Change and Sustainability Institute, Faculdade de Ciências, Universidade de Lisboa , Campo Grande, Lisboa , Portugal
4. Institute of Ecology and Evolution, University of Bern , Bern , Switzerland
5. Swiss Institute of Bioinformatics , Lausanne , Switzerland