1. Biology Department, University of Iowa , Iowa City, IA , United States
2. Institute of Evolutionary Biology, University of Edinburgh , Edinburgh, Scotland , UUnited Kingdom
3. Biological Sciences, Binghamton University , Binghamton, NY , United States
4. Biology Department, Farmingdale State College—State University of New York , Farmingdale, NY , United States
5. Department of Ecology and Evolution, University of Chicago , Chicago, IL , United States