1. Department of Oral Anatomy and Cell Biology, Graduate School of Medical and Dental Sciences, Kagoshima University , Kagoshima , Japan
2. Department of Oral and Maxillofacial Surgery, Graduate School of Medical and Dental Sciences, Kagoshima University , Kagoshima , Japan
3. Department of Anthropology, National Museum of Nature and Science , Tsukuba , Japan
4. Institute of Biotechnology, University of Helsinki , Helsinki , Finland
5. Geological Survey of Finland , Espoo , Finland