1. Université Clermont Auvergne, EA 4678 CIDAM, 63000 Clermont-Ferrand, France (previous address)
2. Bioinformatics Group, Department of Computer Science, University of Freiburg, 79110 Freiburg, Germany
3. Université Clermont Auvergne, INRA, MEDIS, 63000 Clermont-Ferrand, France
4. Université Clermont Auvergne, CNRS, LMGE, 63000 Clermont–Ferrand, France
5. Université Clermont Auvergne, CNRS, LIMOS, 63000 Clermont–Ferrand, France
6. Department of Bioinformatics, Erasmus University Medical Center, Rotterdam, 3015 CE, Netherlands