1. NIHR Health Protection Research Unit in Gastrointestinal Infections, Norwich Medical School, University of East Anglia, Norwich Research Park, Norwich, NR4 7TJ, UK
2. Department of Infection Biology, Institute of Infection and Global Health, Faculty of Health and Life Sciences, University of Liverpool, Liverpool Science Park IC2, 146 Brownlow Hill, Liverpool, L3 5RF, UK
3. Comparative Biomedical Sciences, Royal Veterinary College, Royal College Street, University of London, London, NW1 0TU, UK
4. Department of Science and Technology, Faculty of Health and Science, James Hehir Building, Neptune Quay, University of Suffolk, Ipswich, IP4 1QJ, UK
5. School of Pharmacy and Biomolecular Sciences, Liverpool John Moores University, Liverpool, L3 3AF, UK
6. Faculty of Natural Sciences, Keele University, Staffordshire, ST5 5BG, UK