1. National Engineering Laboratory for Animal Breeding; Key Laboratory of Animal Genetics, Breeding, and Reproduction, Ministry of Agriculture; College of Animal Science and Technology, China Agricultural University, Beijing, 100193, China
2. Department of Agricultural, Food & Nutritional Science, University of Alberta, Edmonton, T6G 2C8, Canada
3. Animal Genomics and Improvement Laboratory, BARC, USDA-ARS, Beltsville, MD 20705-2350, USA
4. Meat Animal Research Center, USDA-ARS, Clay Center, NE 68933 USA
5. School of Animal Science, Guizhou University, Guiyang, 550025, China