1. Division of Systems Virology, Department of Microbiology and Immunology, The Institute of Medical Science, The University of Tokyo , Tokyo 1088639 , Japan
2. Institute for Life and Medical Sciences, Kyoto University , Kyoto 6068507 , Japan
3. Graduate School of Pharmaceutical Sciences, Kyoto University , Kyoto 6068501 , Japan
4. Department of AI Systems Medicine, M&D Data Science Center, Tokyo Medical and Dental University , Tokyo 1138510 , Japan
5. Bioinformatics Laboratory, Faculty of Medicine, University of Tsukuba , Tsukuba 3050821 , Japan
6. Center for Artificial Intelligence Research, University of Tsukuba , Tsukuba 3058577 , Japan
7. Laboratory of Systems Genomics, Graduate School of Frontier Sciences, The University of Tokyo , Kashiwa 2778561 , Japan
8. International Research Center for Infectious Diseases, The Institute of Medical Science, The University of Tokyo , Tokyo 1088639 , Japan
9. International Vaccine Design Center, The Institute of Medical Science, The University of Tokyo , Tokyo 1088639 , Japan
10. Graduate School of Medicine, The University of Tokyo , Tokyo 1130033 , Japan
11. Graduate School of Frontier Sciences, The University of Tokyo , Kashiwa 2778561 , Japan
12. Collaboration Unit for Infection, Joint Research Center for Human Retrovirus infection, Kumamoto University , Kumamoto 8600811 , Japan
13. CREST, Japan Science and Technology Agency , Kawaguchi 3320012 , Japan