1. Department of Crop and Soil Sciences, Washington State University, 1170 NE Stadium Way, Pullman, Washington, 99164-6420, USA
2. Key Laboratory of Agricultural Animal Genetics, Breeding and Reproduction, Ministry of Education, College of Animal Science and Technology, Huazhong Agricultural University, 1 Shizishan Street, Wuhan, Hubei, 430070, China
3. School of Electrical Engineering and Computer Science, Washington State University, 355 NE Spokane Street, Pullman, Washington, 99164-2752, USA