1. Technology Development, Earlham Institute, Norwich Research Park, Norwich NR4 7UZ, UK
2. Department of Cell and Developmental Biology, John Innes Centre, Norwich Research Park, Norwich NR4 7UH, UK
3. Cell and Molcular Sciences, The James Hutton Institute, Invergowrie, Dundee DD2 5DA, UK
4. The New Zealand Institute for Plant & Food Research Limited, Palmerston North 4442, New Zealand
5. Department of Life Sciences, Natural History Museum, Cromwell Road, London WC2 5BD, UK