1. Department of Evolutionary Biology and Environmental Studies, University of Zurich , Zurich 8057, Switzerland
2. Seychelles Islands Foundation , Victoria, Republic of Seychelles
3. Zoological Museum, University of Zurich , Zurich 8006, Switzerland
4. Indian Ocean Tortoise Alliance , Ile Cerf, Victoria, Republic of Seychelles
5. Zoo Zürich , Zurich 8044, Switzerland
6. Centre for Ecology and Conservation, College of Life and Environmental Sciences, University of Exeter , Penryn, Cornwall, TR10 9FE, UK
7. Institute of Biology, University of Neuchâtel , Neuchâtel 2000, Switzerland