1. Department of Cell Biology, Van Andel Institute , 333 Bostwick NE, Grand Rapids, MI 49503 , USA
2. Department of Obstetrics, Gynecology and Reproductive Biology, College of Human Medicine, Michigan State University , Grand Rapids, MI , USA
3. Department of Pathobiology and Diagnostic Investigation, Veterinary Diagnostic Laboratory, College of Veterinary Medicine, Michigan State University , East Lansing, MI , USA