1. Northeast Forestry University College of Wildlife and Protected Areas, , Harbin 150040, China
2. Chinese Academy of Sciences Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, , Beijing 100101, China
3. Xinjiang Northwest Star Information Technology Co., Ltd. , Urumqi 830000, China
4. State Forestry Administration Key Laboratory of Conservation Biology, , Harbin 150040, China