1. McCaig Institute for Bone and Joint Health , Cumming School of Medicine, , Calgary, AB T2N 4Z6 , Canada
2. University of Calgary , Cumming School of Medicine, , Calgary, AB T2N 4Z6 , Canada
3. Department of Cell Biology and Anatomy , Cumming School of Medicine, , Calgary, AB T2N 4N1 , Canada
4. Alberta Children’s Hospital Research Institute, University of Calgary , Cumming School of Medicine, , Calgary, AB T2N 4N1 , Canada