1. Cell and Information, PRESTO, Research Development Corporation of Japan (JRDC), and Department of Physiology, Toho University School of Medicine Tokyo 143
2. Department of Life Science, Faculty of Bioscience and Biotechnology, Tokyo Institute of Technology Yokohama 227
3. Department of Ophthalmology, Toho University School of Medicine Tokyo 105, Japan
4. Department of Physiology and Anatomy, Kyoritsu College of Pharmacy Tokyo 105, Japan