1. Telomere Biology Laboratory, Cancer Research UK, London Research Institute, London, WC2A 3LY, UK
2. Cell Cycle Lab, The Francis Crick Institute, 1 Midland Road, London NW1 1AT, UK
3. Bioinformatics and Biostatistics, The Francis Crick Institute, 1 Midland Road, London NW1 1AT, UK
4. Howard Hughes Medical Institute, Department of Chemistry and Biochemistry, BioFrontiers Institute, University of Colorado, Boulder, CO 80309, USA
5. Telomere Biology Section, Laboratory of Biochemistry and Molecular Biology, National Cancer Institute, National Institutes of Health, Bethesda, MD 20892, USA