1. Frazer Institute, Faculty of Medicine, University of Queensland , Brisbane, QLD, Australia
2. Institute for Molecular Bioscience, University of Queensland , Brisbane, QLD, Australia
3. Medical Research Council Integrative Epidemiology Unit, University of Bristol , Bristol, UK
4. K.G. Jebsen Center for Genetic Epidemiology, Department of Public Health and Nursing, Norwegian University of Science and Technology , Trondheim, Norway