1. Department of Biology, Utah State University, 5305 Old Main Hill, Logan, UT 84322, USA
2. Pollinating Insect—Biology, Management, and Systematics Research Unit, U.S. Department of Agriculture—Agricultural Research Service, 1410 North 800 East, Logan, UT 84341, USA
3. Washington State Department of Agriculture, 1111 Washington Street SE, Olympia, WA 98504, USA