1. Department of Statistical Genetics, Osaka University Graduate School of Medicine , Suita , 565-0871, Japan
2. Laboratory of Statistical Immunology, Immunology Frontier Research Center (WPI-IFReC), Osaka University , Suita, 565-0871, Japan
3. Department of Genome Informatics, Graduate School of Medicine, the University of Tokyo , Tokyo , 113-0033, Japan
4. Department of Respiratory Medicine and Clinical Immunology, Osaka University Graduate School of Medicine , Suita, 565-0871 , Japan
5. Department of Infectious Diseases, Keio University School of Medicine , Tokyo, 160-8582 , Japan
6. M&D Data Science Center, Tokyo Medical and Dental University , Tokyo, 113-8510 , Japan
7. Integrated Frontier Research for Medical Science Division, Institute for Open and Transdisciplinary Research Initiatives, Osaka University , Suita, 565-0871 , Japan
8. Department of Immunopathology, Immunology Frontier Research Center (WPI-IFReC), Osaka University , Suita, 565-0871 , Japan
9. Center for Infectious Disease Education and Research (CiDER), Osaka University , Suita, 565-0871 , Japan
10. Department of Respiratory Medicine, Nagoya University Graduate School of Medicine , 65 tsurumai, Showa-ku, Nagoya, 466-8550 , Japan
11. Health Science Research and Development Center (HeRD), Tokyo Medical and Dental University , Tokyo, 113-8510 , Japan
12. Institute of Research, Tokyo Medical and Dental University , Tokyo, 113-8510 , Japan
13. Division of Health Medical Intelligence, Human Genome Center, the Institute of Medical Science, the University of Tokyo , Tokyo, 108-8639 , Japan
14. Department of Pathology and Tumor Biology, Kyoto University , Kyoto, 606-8315 , Japan
15. Institute for the Advanced Study of Human Biology (WPI-ASHBi), Kyoto University , Kyoto, 606-8303 , Japan
16. Department of Medicine, Center for Hematology and Regenerative Medicine, Karolinska Institute , Stockholm, 171 77 , Sweden
17. Division of Gastroenterology and Hepatology, Department of Medicine, Keio University School of Medicine , Tokyo, 160-8582 , Japan
18. Division of Pulmonary Medicine, Department of Medicine, Keio University School of Medicine , Tokyo, 160-8582 , Japan
19. Laboratory for Systems Genetics, RIKEN Center for Integrative Medical Sciences , Yokohama, 230-0045 , Japan