1. Department of Integrative Biology, University of Guelph , Guelph, Ontario , Canada
2. Department of Psychology, Neuroscience & Behaviour, McMaster University , Hamilton, Ontario , Canada
3. Department of Evolution, Ecology, and Organismal Biology, The Ohio State University , Columbus, OH , USA
4. Department of Biology, Millersville University , Millersville, PA , USA
5. School of Earth, Atmospheric and Life Sciences, University of Wollongong , Wollongong, New South Wales , Australia
6. Department of Mathematics, The Ohio State University , Columbus, OH , USA