1. Department of Evolutionary Studies of Biosystems, SOKENDAI (The Graduate University for Advanced Studies), Hayama, Kanagawa, Japan
2. Department of Dermatology, Graduate School of Medicine, University of the Ryukyus, Nishihara, Okinawa, Japan
3. Graduate School of Tourism Sciences, University of the Ryukyus, Nishihara, Okinawa, Japan
4. Institute of Primate Research, National Museum of Kenya, Karen, Nairobi, Kenya
5. Molecular Biology Section, Department of Cellular and Molecular Biology, Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan