1. Department of Natural Resources and the Environment, University of Connecticut, Storrs, CT 06269, USA
2. Center for Environmental Sciences and Engineering, University of Connecticut, Storrs, CT 06269, USA
3. Department of Bioscience, Arctic Research Centre, Aarhus University, Roskilde, DK-4000, Denmark
4. Great Lakes Institute for Environmental Research, University of Windsor, Windsor, ON N9B 3P4, Canada
5. Department of Biology, Norwegian University of Science and Technology, Trondheim, NO-7491, Norway
6. Wildlife and Landscape Science Directorate, Science and Technology Branch, Environment Canada, National Wildlife Research Centre, Carleton University, Ottawa, ON K1A 0H3, Canada