1. Department of Wine and Food Science, University of Adelaide, PMB 1 Glen Osmond, SA 5064, Australia
2. Institute for Wine Biotechnology, Department of Oenology and Viticulture, Private Bag X1, Stellenbosch University, Stellenbosch 7600, South Africa
3. Division of Biological Science, Graduate School of Science and Technology, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma, Nara 630-0192, Japan