1. State Key Laboratory of Bioreactor Engineering, East China University of Science and Technology , Shanghai 200237, China
2. Department of Biology and Biological Engineering, Chalmers University of Technology , Gothenburg SE41296 , Sweden
3. BioInnovation Institute , Ole Maaløes Vej 3, DK2200 Copenhagen N, Denmark
4. Key Laboratory of Systems Microbial Biotechnology, Tianjin Institute of Industrial Biotechnology , Chinese Academy of Sciences, Tianjin 300308 , China