1. Graduate School of Life Sciences, Tohoku University , Sendai, 980-8577 Japan
2. Faculty of Agriculture, Kyushu University , Fukuoka, 812-8581 Japan
3. PRESTO, JST , Saitama, 332-0012 Japan
4. Division of Biological Sciences, Faculty of Science, Hokkaido University , Sapporo, 060-0810 Japan
5. Institute for Chemical Research, Kyoto University , Uji, 611-0011 Japan
6. Key Laboratory of Ecology and Environmental Science in Guangdong Higher Education, School of Life Sciences, South China Normal University , Guangzhou 510631, China