1. CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences , Kunming, Yunnan 650201 , China
2. Yunnan Key Laboratory for Integrative Conservation of Plant Species with Extremely Small Population, Kunming Institute of Botany, Chinese Academy of Sciences , Kunming, Yunnan 650201 , China
3. School of Integrative Plant Science, Section of Plant Biology and the L. H. Bailey Hortorium, Cornell University , Ithaca, NY 14850 , USA
4. BTI Computational Biology Center, Boyce Thompson Institute , Ithaca, NY 14853 , USA
5. University of Chinese Academy of Sciences , Beijing 100049 , China
6. Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences & Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences , Mengla, Yunnan 666303 , China
7. State Key Laboratory of Phytochemistry and Plant Resources in West China, Kunming Institute of Botany, Chinese Academy of Sciences , Kunming, Yunnan 650201 , China