1. Key Laboratory of Southern Subtropical Plant Diversity, Fairy Lake Botanical Garden, Shenzhen & Chinese Academy of Sciences , Shenzhen, 518004 , China
2. College of Life Sciences, South China Agricultural University , Guangzhou, 510642 , China
3. Global Biodiversity Conservancy 144/124 Moo 3 , Soi Bua Thong, Bangsalae, Sattahip, Chonburi 20250 , Thailand
4. New York Botanical Garden , Bronx , NY 10458 , USA