1. Key Laboratory of Plant Resources Conservation and Sustainable Utilization/Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden, Chinese Academy of Sciences , Guangzhou, 510650 , China
2. University of Leicester, Institute for Environmental Futures, Department of Genetics and Genome Biology , Leicester, LE1 7RH , UK
3. South China National Botanical Garden, Chinese Academy of Sciences , Guangzhou, 510650 , China
4. Center of Conservation Biology, Core Botanical Garden, Chinese Academy of Sciences , Guangzhou , China