1. Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences , Menglun, Yunnan , China
2. University of Chinese Academy of Sciences , Beijing , China
3. Institute of Ecology and Environmental Sciences, CNRS UMR 7618, Sorbonne University , Paris , France
4. Center of Conservation Biology, Core Botanical Gardens, Chinese Academy of Sciences , Menglun, Yunnan , China