1. Division of Applied Life Sciences, Graduate School of Agriculture, Kyoto University, Kitashirakawa-oiwakecho, Sakyo-ku, Kyoto, Japan
2. Laboratory of Industrial Microbiology, Graduate School of Agriculture, Kyoto University, Kitashirakawa-Oiwakecho, Sakyo-ku, Kyoto, Japan
3. Research Institute for Bioscience Products and Fine Chemicals, AJINOMOTO CO., INC., Kawasaki, Kanagawa, Japan
4. Biotechnology Research Center and Department of Biotechnology, Toyama Prefectural University, Imizu, Toyama, Japan
5. Department of Materials Chemistry and Bioengineering, National Institute of Technology, Oyama College, Oyama, Tochigi, Japan