1. Division of Applied Life Sciences, Graduate School of Agriculture, Kyoto University , Kitashirakawa Oiwake-cho, Sakyo-ku, Kyoto , Japan
2. Department of RNA Biology and Neuroscience, Osaka University Graduate School of Medicine , 2-2 Yamada-oka, Suita, Osaka , Japan
3. Department of Ophthalmology, Yamaguchi University Graduate School of Medicine , 1-1-1 Minami-Kogushi, Ube, Yamaguchi , Japan
4. Department of Applied Biological Chemistry, Graduate School of Agriculture, Osaka Metropolitan University , 1-1 Gakuen-cho, Naka-ku, Sakai, Osaka , Japan
5. Department of Biological Chemistry, Graduate School of Science, Osaka Metropolitan University , 1-2 Gakuen-cho, Naka-ku, Sakai, Osaka , Japan