1. Department of Food science, National Fisheries University , Shimonoseki , Japan
2. Department of Biological Chemistry, Faculty of Agriculture, Yamaguchi University , Yamaguchi , Japan
3. Department of Applied Bioresource Science, Faculty of Agriculture, Ehime University , Matsuyama , Japan
4. Research and Development Center for Industrial Fermentation (CINDEFI; UNLP, CONICET La Plata), School of Science, La Plata National University , 47 y 115 (B1900ASH), La Plata , Argentina