1. Plymouth Marine Laboratory, Prospect Place, West Hoe, Plymouth PL1 3DH, UK
2. School of Environmental Sciences, University of Liverpool, Liverpool L69 3GP, UK
3. Centre for Ocean Life, National Institute for Aquatic Resources, Technical University of Denmark, Kavalergarden 6, Charlottenlund 2920, Denmark
4. Sweet Briar, Totnes TQ9 5LH, UK
5. College of Life and Environmental Sciences, University of Exeter, Hatherly Laboratories, Prince of Wales Road, Exeter EX4 4PS, UK