1. CAS Key Laboratory of Marine Ecology and Environmental Sciences , Institute of Oceanology, , 7 Nanhai Rd, Shinan District, Qingdao, Shandong Province 266071 , P. R. China
2. Chinese Academy of Sciences , Institute of Oceanology, , 7 Nanhai Rd, Shinan District, Qingdao, Shandong Province 266071 , P. R. China
3. Laboratory for Marine Ecology and Environmental Science , Qingdao Marine Science and Technology Center, 168 Wenhai Middle Rd, Jimo District, Qingdao, Shandong Province 266237 , P. R. China
4. Center for Ocean Mega-Science, Chinese Academy of Sciences , 7 Nanhai Rd, Shinan District, Qingdao, Shandong Province 266071 , P. R. China
5. University of Chinese Academy of Sciences , 1 Yanqihu East Rd, Huairou District, Beijing 101408 , P. R. China
6. Key Laboratory of Marine Ecosystem Dynamics, Second Institute of Oceanography, Ministry of Natural Resources , 36 Baochubei Rd, Xihu District, Hangzou, Zhejiang Province 310012 , P. R. China
7. Key Laboratory of Nearshore Engineering Environment and Ecological Security of Zhejiang Province , 36 Baochubei Rd, Xihu District, Hangzou, Zhejiang Province 310012 , P. R. China
8. Aix-Marseille University , Toulon University, CNRS, IRD, , 163 Avenue de Luminy - Bâtiment Méditerranée, 13288 Marseille cedex 09 , France
9. Mediterranean Institute of Oceanography UM110 , Toulon University, CNRS, IRD, , 163 Avenue de Luminy - Bâtiment Méditerranée, 13288 Marseille cedex 09 , France