1. Department of Arctic and Marine Biology , Faculty of Biosciences, Fisheries and Economics, , Framstredet 39, 9019 Tromsø, Norway
2. UiT The Arctic University of Norway , Faculty of Biosciences, Fisheries and Economics, , Framstredet 39, 9019 Tromsø, Norway
3. Department of Arctic Biology, The University Centre in Svalbard , PO Box 156, N-9171 Longyearbyen, Svalbard, Norway
4. Department of Forestry and Wildlife Management, Inland Norway University of Applied Sciences , PO Box 400 Vestad, 2418 Elverum, Norway
5. Norwegian College of Fishery Science , UiT The Arctic University of Norway, Muninbakken 21, 9037 Tromsø, Norway