1. Department of Biological Sciences, MQ Marine Research Centre, Macquarie University, North Ryde, NSW 2109, Australia
2. Antarctic Climate & Ecosystems CRC, University of Tasmania, Hobart, TAS 7001, Australia
3. CSIRO Oceans and Atmosphere, Castray Esplanade, Hobart, TAS 7000, Australia
4. Department of Geology, Colgate University, Hamilton, NY 13346, USA