1. Institute of Oceanology, Chinese Academy of Sciences CAS Key Laboratory of Marine Ecology and Environmental Sciences, , 7 Nanhai Road, Qingdao 266071 , China
2. Laboratory for Marine Ecology and Environmental Science, Pilot National Laboratory for Marine Science and Technology (Qingdao) , 1 Wenhai Road, Qingdao 266237 , China
3. Chinese Academy of Sciences Center for Ocean Mega-Science, , 7 Nanhai Road, Qingdao 266071 , China