1. CREAF , Cerdanyola del Vallès, Catalonia E-08193 , Spain
2. Department of Bioscience, Aarhus University , Aarhus C DK-8000 , Denmark
3. CEAB, CSIC , Blanes, Catalonia E-17300 , Spain
4. Department of Environmental and Biological Sciences, University of Eastern Finland , Kuopio FI-70211 , Finland
5. área de Biodiversidad, Gestión ambiental de Navarra-Nafarroako Ingurumenkudeaketa (GAN-NIK) , Pamplona-Iruñea, Navarra E-31001 , Spain
6. Departament de Biologia Evolutiva, Ecologia i Ciències Ambientals, Universitat de Barcelona , Barcelona, Catalonia E-08028 , Spain
7. CSIC , Bellaterra E-08193 , Spain