1. Laboratory of Reproductive and Developmental Biology , Department of Biological Sciences, Faculty of Science, , Sapporo , Japan
2. Hokkaido University , Department of Biological Sciences, Faculty of Science, , Sapporo , Japan
3. Department of Molecular Biology and Biotechnology , Faculty of Science, , Peradeniya , Sri Lanka
4. University of Peradeniya , Faculty of Science, , Peradeniya , Sri Lanka